The Bottom lane has five towns, spaced evenly along the lane. See The Map. The last player to move a unit past a town owns it. Owning a town lets you spend land on its upgrade. Click a town you own to start or stop that research. Two towns of the same upgrade pay into the same research at once. Land income is on Gold, Land, and Banks.
From either end the towns are Defense, Speed, Damage, Speed, Defense. Research spends 1 land per second until the rank is paid for. Each rank costs the same land price. There are five ranks.
Ranks of the same upgrade add together. Rank 3 Damage is one +30% attacker modifier, not three separate ×1.1 multipliers. Rank 3 Defense is one ×0.7 incoming factor. Defense then multiplies with cover and other defensive factors. How those modifiers combine is on Damage.
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